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Secondly, according to the stem and leaf anatomy, leaf epidermis, pollen morphology and floral development, it is found that there are no close relations between Acorus and Typhales, especially in the aspect of floral development. In the course of floral development of A. calamus, neither bracteloe primordium nor common primordium occurs in the whole course. Early development of the flower is strongly unidirectional. Three shield carpels develop into a pistil. In Sparganium fallax, each female flower has a bracteole. The bracteole primordium and the floral primordium originate from a complex structure. A single saccate carpel develops into a pistil. In staminate flowers, petal and stamen primordia initiate spirally.

菖蒲属与香蒲目的茎叶解剖特征、叶表皮特征、花粉特征及花器官发生特点等亦存在着明显区别,特别是花器官发生特点差异尤为显著,如菖蒲属花器官发生过程中,没有苞片原基,也不存在任何形式的复合原基,花被与雄蕊原基均以单向性方式发生,雌蕊由3心皮发育而成,具有"盾状心皮"的特点等,黑三棱属则有苞片原基发生,且苞片原基与雌花原基起源于共同的复合结构,雄花的花被原基和雄蕊原基均以螺旋状方式发生,雌蕊由单心皮发育而成,未发现有2或3个心皮原基或其痕迹的现象,体现出"囊状心皮"的特点。

The lack in Galitzkya of simple and forked trichomes and staminal appendages, and the development of thick, branched caudices and saccate, lateral sepals clearly distinguish it from Berteroa .

在单和分叉的毛状体和雄蕊的附属物的Galitzkya里的缺乏,后面属有单和分叉的毛状体,雄蕊的附属物和无气囊的萼片。

Flowers small to medium-sized, bracteate and often bracteolate, bisexual or unisexual, actinomorphic or rarely zygomorphic, hypogynous to epigynous, floral cup shallow to urn-shaped, sometimes absent; sepals 4 or 5(–10), sometimes absent, imbricate, usually persistent; petals absent or 4 or 5, yellow, white, greenish or red, often ribbonlike and circinate in bud, caducous; stamens 4, 5, or many, free, rarely arranged in 2 whorls with the inner whorl staminodal, development of polyandrous androecia centripetal or centrifugal; anthers basifixed, thecae mostly bisporangiate, each opening by two valves or a simple longitudinal slit, or monosporangiate and opening by a single valve (Exbucklandia, Hamamelis and the genera of the S hemisphere), connective protruding; disk scales sometimes present between stamens and carpels.

花小对中型,具苞片和通常,萼片4或5(-10),有时无,覆瓦状,通常宿存;花瓣无或者4-5,黄色,白色,带绿色或者红,通常在芽期带状的并且拳卷,早落;雄蕊4,5,或者很多,离生,很少2轮生并且退化雄蕊内部轮生,多雄蕊的雄蕊群向心或离花药基着大多为的囊bisporangiate,以二裂爿或一单纵裂缝每开阔的,花盘鳞片有时片在雄蕊和心皮之间存在。

Sepals 3(or 5); lateral sepals free or connate, margins entire or serrate; lower sepal large, petaloid, usually navicular, funnelform, saccate, or cornute, tapering or abruptly constricted into a nectariferous spur broadly or narrowly filiform, straight, curved, incurved, or ± coiled, swollen at tip, or pointed, rarely 2-lobed, rarely without spur. Petals 5, free, upper petal flat or cucullate, small or large, often crested abaxially, lateral petals free or united in pairs. Stamens 5, alternating with petals, connate or nearly so into a ring surrounding ovary and stigma, falling off in one piece before stigma ripens; filaments short, flat with a scalelike appendage inside; anthers 2-celled, connivent, opening by a slit or pore.

萼片3(或5);侧萼片离生或者合生,边缘全缘或者有锯齿;更低的萼片大,瓣状,通常舟形,漏斗状,囊状,或具角状突起,渐减突然或缢缩进一分泌蜜汁具距或宽狭丝状,花瓣5,离生,上面的花瓣平坦的或者兜状,雄蕊5,与花瓣互生,合生或者近合生成环而围绕子房和柱头,在柱头成熟之前完整地从上面落下;花丝短,扁平,在里面的有一鳞片状附属物;花药2室,靠合,通过一条裂缝或者孔开裂。

The floral primordium is initially circular, along with the edge of which slight concaves occur. The outmost whorl of tepal primordia arises as a result of apical meristem division and the floral primordium then is in the shape of triangle. Two further whorls of tepal primordia arise in spiral acropetal sequence and are alternately arranged. When the innermost whorl of tepal primordia is at the late developmental stage, the foremost stamen primordia arise almost simultaneously in spiral acropetal succession and all of the stamen primordia are ultimately arranged in a triangle-cone form. Late in stamen primordium development, carpel primordia arise as same as the young stamen primordia and grow laterally. Grooves are present on the adaxial side of each carpel and develop into ventral sutures gradually closing up.

观光木的花原基最初为近圆形,随着顶端分生组织的活动,花原基边缘处出现浅凹,形成第一轮花被片原基,此时,花原基呈三角形排列,后两轮花被片原基依次发生,与前一轮互生;在内轮花被片发生的后期,最初几枚雄蕊原基几乎同时出现,呈螺旋状向顶发生,最后排列成三角圆锥状;雄蕊原基发育后期,心皮原基开始发育,形状与发育初期的雄蕊原基相似,随后心皮原基进行侧向生长,在近轴面出现浅凹,进而发育为凹槽,形成腹缝线,最后腹缝线完全愈合。

Stamens numerous, rarely few, free; filaments linear or filiform; anthers latrorse, introrse, or extrorse; sometimes some sterile stamens becoming staminodes.

雄蕊多,有的很少,离生;花丝线形或者丝状;花药侧向纵裂的,向内,或者外向;有时一些不育的雄蕊成为退化雄蕊。

Secondly, according to the stem and leaf anatomy, leaf epidermis, pollen morphology and floral development, it is found that there are no close relations between Acorus and Typhales, especially in the aspect of floral development. In the course of floral development of A. calamus, neither bracteloe primordium nor common primordium occurs in the whole course. Early development of the flower is strongly unidirectional. Three shield carpels develop into a pistil. In Sparganium fallax, each female flower has a bracteole. The bracteole primordium and the floral primordium originate from a complex structure. A single saccate carpel develops into a pistil. In staminate flowers, petal and stamen primordia initiate spirally.

菖蒲属与香蒲目的茎叶解剖特征、叶表皮特征、花粉特征及花器官发生特点等亦存在着明显区别,特别是花器官发生特点差异尤为显著,如菖蒲属花器官发生过程中,没有苞片原基,也不存在任何形式的复合原基,花被与雄蕊原基均以单向性方式发生,雌蕊由3心皮发育而成,具有&盾状心皮&的特点等,黑三棱属则有苞片原基发生,且苞片原基与雌花原基起源于共同的复合结构,雄花的花被原基和雄蕊原基均以螺旋状方式发生,雌蕊由单心皮发育而成,未发现有2或3个心皮原基或其痕迹的现象,体现出&囊状心皮&的特点。

Androgynophore usually present; filaments usually connate into a single tube; staminodes 5, tonguelike or filiform, opposite to sepals, sometimes lacking; anthers 2-celled, longitudinally dehiscent.

雌雄蕊柄通常宿存;花丝通常合生成一单个的管;退化雄蕊5,舌状或者丝状,与萼片对生,有时无;花药2室,纵向开裂。

Stamens 15 in 5 groups of 3; filaments fused to ovary stipe to form androgynophore, upper part widening to enclose ovary; anthers sessile, usually in very congested head, 2-celled; staminodes 5, toothlike, minute.

雄蕊15枚成5 组每组3枚;花丝愈合到子房果柄形成雌雄蕊柄,上半部分加宽成包围子房;无柄的花药,通常成密集头状花序,2室;退化雄蕊5,齿状,小。

Petals white, ovate-elliptic, shallowly boat-shaped, ca. 4 mm. Stamens 6; fertile stamens 3, anterior; staminodes 3, with shorter filaments.

花瓣白色,卵状椭圆形,浅舟状,约4毫米雄蕊6;能育的雄蕊3,前面;退化雄蕊3,带有更短的花丝。

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